In einer postapokalyptischen Welt hat die Menschheit »Nanomachines« entwickelt – künstliche Superzellen, die menschliche Fähigkeiten steigern und Krankheiten heilen können. Verbrecher missbrauchen die Nanomachines jedoch als Drogen. Nur eine Spezialeinheit der Polizei, bestehend aus Polizisten und sogenannten »Hangern« – ehemaligen Süchtigen, die ihre Strafe durch Verbrecherjagd abbüßen – kann sich ihnen entgegenstellen. Als der Polizist Hajime in diese Einheit versetzt wird, bekommt er den Hanger Zeroichi zugeteilt, einen unkooperativen und mürrischen jungen Mann, dem eine Strafe von 512 Jahren auferlegt wurde ...
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